Billy Twelvetrees will make his first start of the Autumn international series against Australia on Saturday.
The Gloucester captain, from Wisborough Green, replaces Owen Farrell at centre for the weekend’s Test at Twickenham
Former Horsham, Pulborough and Haywards Heath colt Twelvetrees, 26, impressed as a second-half replacement in the 28-9 victory over Samoa after being left out of the squad for the defeats against South Africa and New Zealand.
Farrell has been overlooked for the starting XV for the first time since June 2012 after under-performing against New Zealand, South Africa and Samoa .
The experiment of playing the 23-year-old at 12 has been abandoned after only one match with Twelvetrees chosen to line up outside fly-half George Ford.
England head coach Stuart Lancaster said: “It’s good to be able to give Billy his first start of the series.
“He took on board the feedback we gave him when he first came into camp and he has worked hard both with us and at Gloucester. He deserves his chance.
“Owen is a key part of our squad and I’m sure he’ll make a positive contribution should he get the chance.”
England: Mike Brown (Harlequins), Anthony Watson (Bath Rugby), Brad Barritt (Saracens), Billy Twelvetrees (Gloucester Rugby) Jonny May (Gloucester Rugby), George Ford (Bath Rugby), Ben Youngs (Leicester Tigers); Joe Marler (Harlequins), Dylan Hartley (Northampton Saints), David Wilson (Bath Rugby), Dave Attwood (Bath Rugby), Courtney Lawes (Northampton Saints), Tom Wood (Northampton Saints), Chris Robshaw (Harlequins, capt), Ben Morgan (Gloucester Rugby).
Replacements: Rob Webber (Bath Rugby), Matt Mullan (Wasps), Kieran Brookes (Newcastle Falcons), George Kruis (Saracens), James Haskell (Wasps), Richard Wigglesworth (Saracens), Owen Farrell (Saracens), Marland Yarde (Harlequins).